Phiangphak Sukkharak

Phiangphak Sukkharak

Department of Biology, Faculty of Science, Burapha University, Chonburi, 20131, Thailand


I am an associate professor at Burapha University in Thailand. I completed my primary and secondary schools in Nan, where I was born in 1982. Later, I continued my high school and Bachelor’s degree in Chiang Mai because I obtained the Development and Promotion of Science and Technology Talents project scholarship provided by Thai government to support students with a scientific interest to study up to a doctorate degree. My interest in bryophytes started when I joined the Forest Restoration Research Unit of Chiang Mai University’s Biology Department (FORRU-CMU) as a volunteer during my second year of Bachelor’s degree at Chiang Mai University. Because of their beauty, when going into the research nursery, I always looked for bryophytes growing in the germinating area and on the floor. I wanted to know more about this interesting group of plants so I submitted a proposal to the Biodiversity Research and Training program of Thailand and got the financial support to study the bryophytes of Kun Wang community forest in Chiang Mai. I rode a motorbike alone to the Kun Wang community forest of Inthanon mountain, which is the highest mountain in Thailand at 2,565 meters asl to collect the bryophytes. I studied the morphological and anatomical details of the bryophytes and identified the specimens on my own, with the help of Sahut Chantanaorrapint for specimen verification. I also visited the Queen Sirikit Botanical Garden library to borrow books of bryophytes. I was able to complete the project and return more than half of the budget awarded to me. Although there were difficulties encountered during the study, I found out that the more I studied bryophytes, the more I loved them and would like to know more. I, therefore, continue studying the liverworts of Khao Nan in Nakhon Si Thammarat for my master’s degree thesis at Chulalongkorn University in Bangkok. In 2007, I met Robbert Gradstein at the World Conference of Bryology in Kuala Lumpur, Malaysia and was invited to Göttingen, Germany to conduct a monograph of the liverwort genus Thysananthus for my PhD thesis under his guidance. After graduation, I wrote a monograph on Mastigolejeunea whereby intermediate taxa between Thysananthus and Mastigolejeunea were found. This lead to the treatment of Mastigolejeunea as a subgenus of Thysananthus based on morphological and molecular evidence. Recently, I have been working on a revision of the liverwort genera of Thailand. The genera Frullania, Pleurozia, and Metzgeria in Thailand are revised. Porella and Acrobulbus are updated and Syzygiella is added to the Thai liverwort record. I am also working on Thysananthus and Frullania for the Flora of Singapore project. I currently serve as a member of the Nomenclature Committee for Bryophytes of the International Association of Plant Taxonomists(2017-2023)andthe Editorial Board of Journal of the Hattori Botanical Laboratory (2017-present). Moreover, after receiving the Young Scientist Award from the Foundation for the Promotion of Science and Technology under the Patronage of His Majesty the King in 2015 until now, I have been invited to give stimulating talks and to conduct the outreach activities for students not only in universities but also high schools throughout Thailand. My aim is to share my research experience, encourage and motivate them into scientific careers, as well as increase their awareness of this important and beautiful, yet understudied group of plants. As a woman studying a little known group of plants in Thailand, I am proud of what I do. I have never felt advantages, disadvantages, or regret because I have a passion for bryophytes and I believe in my ability and perseverance that keeps me doing and accomplishing anything I set my mind to.

Rxng ???s?tr?c?ry? dr.Phe?yng ph?ktr? s?uk?h r?ks??? p?h?kh wich? ch?wwithy? kh?a withy????s?tr? mh??withy?l?y b?rph? x?mel?: Ppsukk@gmail.Com, phiangphak@buu.Ac.Th p?ccub?n d?rng t?h?æ?ng rxng ???s?tr?c?ry? ? mh??withy?l?y b?rph? s??r?c k?r ???ks??? n? rad?b prat?hm ???ks??? læa m??hym???ks??? txn t?n c?k c?ngh?w?d ??n t?x m? d?? ???ks??? n? rad?b m??hym???ks??? txn pl?y th?? rongre?yn yuphr?ch withy?l?y c?ngh?w?d che?yngh??m? læa rad?b pri??? tr? s??k?h? ch?wwithy? kh?a withy????s?tr? mh??withy?l?y che?yngh??m? p?h?y t?? khorngk?r ph?t?hn? læa s??ngs?erim p?h?? m? khw?m s??m?rt?h phi??es?? th?ng withy????s?tr? læa thekhnoloy? (phs?wth.) S??ng p?n khorngk?r th?? s??ngs?erim n?kre?yn th?? s?nc? ke??yw k?b withy????s?tr? læa s?n?bs?nun h??? ???ks??? d??n withy????s?tr? cnt?h?ng rad?b pri??? xek doy khw?m s?nc? ke??yw k?b b?r xo f?t? (bryophytes) reìm t?n rah???ng th?? ???ks??? n? rad?b pri??? tr? ch?n p? th?? 2 k?h?a p?n x?s?? s?m?khr k?hxng h??wy wic?y k?r f???nf? p?? (Forest Restoration Research Unit: FORRU) k?hxng p?h?kh wich? ch?wwithy? kh?a withy????s?tr? mh??withy?l?y che?yngh??m? pher?a khw?m s?wy ng?m k?hxng b?r xo f?t? th?h??? thuk khr?ng th?? dein k?hê?p? n? rongre??xn t?n kl?? ca mxng h?? b?r xo f?t? th?? ceri? xy?? bn ph???nth?? phe?a t?n kl?? læa ph???n k?hxng rongre??xn læa t?xngk?r re?yn r?? ke??yw k?b ph??ch klùm n?? h??? m?k yìng k?h??n c?ng s?enx khorng r??ng ng?n wic?y t?x khorngk?r ph?t?hn? xngkh? khw?m r?? læa ???ks??? nyob?y k?r c?dk?r thr?phy?kr ch?wp?h?ph n? prathe??th?y læa d?? r?b k?r phic?r?? c?ds?rr thun wic?y h??? d?nein k?r wic?y ke??yw k?b khw?m h?l?kh?l?y k?hxng b?r xo f?t? briwe? p?? chum chm b??n k?hun w?ng c?ngh?w?d che?yngh??m? doy d?? k?h?? rt?hc?kry?nynt? k?h??n p? k?b t?w x??ng b?r xo f?t? n? p?? chum chm b??n k?hun w?ng s??ng xy?? ph???nth?? dxy xin thn nth? th?? m? yxd k?he? th?? s??ng th??s?ud n? prathe??th?y ???ks??? l?ks???a th?ng s?????h?n withy? læa k?ywip?h?kh læa rabu chnid k?hxng b?r xo f?t? d?wy tnxeng læa d?? khw?m ch?wyh?el??x c?k rxng ???s?tr?c?ry? dr.S?h??ch c?n thn? xrphinth? n? k?r trwc s?xb khw?m t?h?k t?xng k?hxng k?r rabu chnid t?xng p? th?? s?wn phvks?????s?tr? s?md?c phran?ngcê? s?i riki ti? phe???x y??m h?n?ngs???x th?? ke??yw k?b b?r xo f?t? m? ???ks??? prakxb k?r th? khorngk?r wic?y cn n? th??s?ud khorngk?r wic?y k? s??r?c læa d?? s??ng kh??n thun wic?y m?kk?? khr??ng h?n??ng k?hxng th?? d?? r?b k?r c?ds?rr m? h??? kæ? khorngk?r ph?t?hn? xngkh? khw?m r?? læa ???ks??? nyob?y k?r c?dk?r thr?phy?kr ch?wp?h?ph n? prathe??th?y rah???ng th?? d?nein khorngk?r wic?y n?? mæ? ca l?b?k tæ? k? phb ?? yìng ???ks??? b?r xo f?t? k? yìng r?k læa xy?k ca ???ks??? m?k k?h??n c?ng d?? k?hê? ???ks??? t?x n? rad?b pri??? tho s??k?h? phvks?????s?tr? kh?a withy????s?tr? cu??lngkr?? mh??withy?l?y doy th? withy?niphn?h? ke??yw k?b khw?m h?l?kh?l?y k?hxng li wexr? weir?t (liverwort) briwe? k?he? n?n c?ngh?w?d nkhr??r??hrrmr?ch læa n? p? ph.??. 2550 D?? phb k?b Robbert Gradstein n? ng?n pra chum b?r xo f?t? withy? n? rad?b lok (World Conference of Bryology) ? me??xng k?wl?l?mpexr? prathe?? m?lese?y læa d?? r?b chei? h??? p? ???ks??? t?x n? rad?b pri??? xek th?? mh?? withy? l?y keiththing ngein (University of Göttingen) prathe?? s?h?ph?n?h? s???h?r?r???h yexrmn? læa th? withy?niphn?h? n? rad?b pri??? xek n? h??wk??x k?r ???ks??? thbthwn li wexr? weir?t s?kul Thysananthus n? rad?b lok doy m? Robbert Gradstein p?n x?c?ry? th?? pr?ks??? withy?niphn?h? h?l?ng s??r?c k?r ???ks??? d?? ???ks??? thbthwn li wexr? weir?t s?kul Mastigolejeunea n? rad?b lok læa ne???xngc?k phb s?m?chik th?? m? l?ks???a th?ng s?????h?n withy? th?? k??k??ng rah???ng s?kul Thysananthus læa s?kul Mastigolejeunea c?ng d?? m? k?r ???ks??? d??n s?????h?n withy? læa khw?m s??mph?n?h? cheing wiw?t?hn?k?r rad?b molekul læa th?h??? s?kul Mastigolejeunea t?h?k ld rad?b h??? p?n s?kul ?xy k?hxng s?kul Thysananthus ng?n wic?y th?? k?l?ng d?nein k?r xy?? n? p?ccub?n kh??x k?r ???ks??? thbthwn li wexr? weir?t n? prathe??th?y doy s?kul th?? d?? ???ks??? thbthwn p? læ?w d??kæ? s?kul Frullania s?kul Pleurozia læa s?kul Metzgeria nxkc?k n?? d?? m? k?r pheìmteim k??xm?l k?hxng s?kul Porella læa Acrobulbus n? prathe??th?y h??? p?n p?ccub?n læa d?? r?yng?n k?r phb s?kul Syzygiella khr?ng ræk n? prathe??th?y x?k th?ng y?ng d?? ???ks??? s?kul Thysananthus læa s?kul Frullania n? khorngk?r Flora of Singapore project k?hxng prathe?? s?ingkhpor? læa th? h?n??th?? p?n krrmk?r k?r t?ng ch???x b?r xo f?t? (Nomenclature Committee for Bryophytes) k?hxng the International Association of Plant Taxonomists (2017-2023) læa p?n krrmk?r kxng brr???hik?r k?hxng w?rs??r Journal of the Hattori Botanical Laboratory (2017-p?ccub?n) nxkc?k n?? h?l?ngc?k th?? d?? r?b r?ngw?l n?k withy????s?tr? rùn h??m? prac? p? ph.??. 2558 C?k m?lni?hi s??ngs?erim withy????s?tr? læa thekhnoloy? n? phrabrm r?ch?pt?h?mp?h? k?d?? r?b chei? h??? brry?y phi??es?? n? ng?n prachum th?ng wich?k?r læa kickrrm th?ng wich?k?r t??ng «s??h?r?b n?kre?yn nis?it n?k???ks??? th??w prathe?? doy m? w?tt?hupras?ngkh? phe???x t?h??ythxd pras?bk?r?? k?r th? wic?y s?r??ng ræng b?nd?l c? n? x?ch?ph n? d??n withy????s?tr? tlxd cn s??ngs?erim h??? trah?n?k t?h?ng khw?m s??kh?? k?hxng b?r xo f?t? th?? m? khw?m s??kh?? doy c?heph?a t?x rabb niwe?? m? khw?m s?wyng?m læa y?ng m? k?r ???ks??? ke??yw k?b ph??ch klùm n?? n?xy doy s??wnt?w r??s??k p?h?mic? th?? p?n p?h??h??ing th?? ???ks??? klùm ph??ch b?r xo f?t? s??ng p?n th?? r??c?k n?xy m?k n? prathe??th?y m?? khey khid ?? p?n k??x d?? per?yb k??x s?e?yper?yb h?r??x t?xthæ? tæ? x??ng d? pher?a r?k læa s?n c? b?r xo f?t? læa che???x n? ???kyp?h?ph læa khw?m mùng m??n t?ngc? k?hxng tnxeng th?? ca th?ng?n wic?y d??n n?? t?x p? x??ng t?x ne???xng læa th? thuk x??ng h??? s??r?c d?? d?ng th?? c? t?xngk?r


Ho, B.-C. & Sukkharak, P. (2021). Thysananthus ciliaris (Lejeuneaceae, Marchantiophyta), a rare species new to Singapore from the Singapore Botanic Gardens. The Gardens’ Bulletin Singapore, 73(2), 237-244.DOI:

Sukkharak, P. (2021). Syzygiella securifolia (Marchantiophyta, Adelanthaceae), a new genus and species record for Thailand. Acta Botanica Hungarica, 63(3-4), 465-470.DOI:

Chantanaorrapint, S., Sevanakun, N., Suwanmala, O., Sukkharak, P.,Printarakul, N. &He, S.(2021). The moss family Rhachitheciaceae (Dicranales, Bryophyta) in Thailand. Phytotaxa, 520(3), 279-284. DOI:

He, S., Sukkharak, P. &Chantanaorrapint, S.(2021). Taxonomic updates and treatments of Hypnum (Hypnaceae, Bryophyta) in Thailand and five new records for Thai moss flora. Hattoria, 12, 27-39.

Feldberg, K., Gradstein, S. R., Gröhn, C., Heinrichs, J., von Konrat, M., Mamontov, Y. S., Renner, M. A. M., Roth, M., Schäfer-Verwimp, A., Sukkharak, P. & Schmidt, A. R. (2021). Checklist of fossil liverworts suitable for calibrating phylogenetic reconstructions. Bryophyte Diversity and Evolution 43(1), 14-71. DOI:

Printarakul, N., Jampeetong, A., Pongkawong, U., Sukkharak, P., Kraichak, E., Adulkittichai, K. & Chantanaorrapint, S. (2021). Koponobryum papillosum Printarakul & Chantanaorr., sp. nov. (Pottiaceae, Bryophyta), a new moss species from northern Thailand. Cryptogamie, Bryologie 42(9),143-148. DOI:

Sukkharak, P. (2021). An account of the genus Acrobolbus Nees (Marchantiophyta: Acrobolbaceae) in Thailand with a new record, Acrobolbus knightii (Mitt.) Briscoe. Phytotaxa, 482(3), 293-296. DOI:

Sukkharak, P. & Chantanaorrapint, S. (2020). The liverwort genus Metzgeria (Metzgeriaceae, Marchantiophyta) in Thailand. Phytotaxa, 441(3), 251-262. DOI: https://doi:10.11646/phytotaxa.441.3.2

Printarakul, N., Sukkharak, P. &Chantanaorrapint, S. (2019). The genus Calycularia (Calyculariaceae, Marchantiophyta)in Thailand. Thai Forest Bulletin, 47(1), 108-112. DOI:

Sukkharak, P. & Chantanaorrapint, S. (2018). An account of the liverwort genus Porella in Thailand with a new record, P. obtusata var. macroloba and the occurrence of asymmetrical underleaves associated with left-right symmetry in the genus. Phytotaxa, 385(2), 77-84. DOI:

Chantanaorrapint, S., Sukkharak, P. & He, S. (2018). Thysananthus ciliaris (Lejeuneaceae, Marchantiophyta): a new record of Thysananthus species for Thailand.Telopea, 21, 153-156. DOI:

He, S., Yan, X.-L., He, L. & Sukkharak, P. (2018). Notes on the distribution of Venturiella perrottetii (Bryophyta: Erpodiaceae), with new records reported from Myanmar and Thailand.Phytotaxa, 362(3), 297-300. DOI:

He, S., Sukkharak, P. & Chantanaorrapint, S. (2018). Pseudoparaphysanthus linisii (Neckeraceae), a new combination and a new record for mainland South-east Asia. Journal of Bryology, 40(2), 143-148. DOI:

Sukkharak, P. (2018). A revision of the genus Frullania (Marchantiophyta: Frullaniaceae) in Thailand. Nova Hedwigia, 106(1-2), 115-207. DOI:

Sukkharak, P. & Gradstein, S. R. (2017). Phylogenetic study of Mastigolejeunea (Marchantiophyta: Lejeuneaceae) and an amended circumscription of the genus Thysananthus. Phytotaxa, 326(2), 91-107. DOI:

Sukkharak, P. (2017). A revision of the genus Pleurozia (Marchantiophyta: Pleuroziaceae) in Thailand and little known morphological features of the genus. Phytotaxa, 309(3), 201-216. DOI:

Sukkharak, P. (2017). An additional variant of Jubula hutchinsiae subsp. javanica (Marchantiophyta: Jubulaceae) in Thailand. Telopea, 20, 105-108. DOI:

Söderström, L., Hagborg, A., von Konrat, M., Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E., Cargill, D. C., Costa, D. P., Crandall-Stotler, B. J., Cooper, E. D., Dauphin, G., Engel, J. J., Feldberg, K., Glenny, D., Gradstein, S. R., He, X., Heinrichs, J., Hentschel, J., Ilkiu-Borges, A. L., Katagiri, T., Konstantinova, N. A., Larraín, J., Long, D. G., Nebel, M., Pócs, T., Puche, F., Reiner-Drehwald, E., Renner, M. A. M., Sass-Gyarmati, A., Schäfer-Verwimp, A., Moragues, J. G. S., Stotler, R. E., Sukkharak, P., Thiers, B. M., Uribe, J., Vá?a, J., Villarreal, J. C., Wigginton, M., Zhang, L. & Zhu, R.-L. (2016). World checklist of hornworts and liverworts. PhytoKeys, 59, 1-828. DOI:

Sukkharak, P. & He, S. (2015). Schwetschkeopsis fabronia (Schwägr.) Broth. (Bryophyta: Fabroniaceae): a species new to Thailand. Telopea, 18, 371-374. DOI:

Parnell, J. A. N., Pilla, F., the Thai Biogeography group (comprising Simpson, D. A., van Welzen, P. C., Chayamarit, K., Chantaranothai, P., Boyce, P. C., Bygrave, P., Byrne, C., Chen, S., Couch, C., Curtis, T., Dransfield, S., Duyfjes, B. E. E., Eianthong, W., Esser, H. J., Grote, P. J., Hua, Z., Jebb, M. H. P., Kirkup, D. W., ke Loc, P., Larsen, S. S., Macklin, J., Madern, A., Meade, C., Merklinger, F., Middleton, D. J., Moat, J., Muasya, A. M., Nakmuenwai, P., Pederson, H., Pendry, C. A., Prajaksood, A., Pooma, R., Preusapan, K., Puglisi, C., Sathapattayanon, A., Sukkharak, P.,Staples, G., Strijk, J., Suddee, S., Sungkaew, S., Tangjitman, K., Teerwatananon, A., Tovaranonte, J., Ung, T., Trias-Blasi, A., de Wilde, W.  J.  J. O., Wilkin, P. & Yahara, T. (2015). A re-examination of the life and work of A.F.G. Kerr and of his colleagues and friends. Thai Forest Bulletin (Botany), 43, 111-131.

Juengprayoon, W., Sukkharak, P. & Chantanaorrapint, S. (2015). Genus Schistochila Dumort. (Schistochilaceae, Marchantiophyta) in Thailand. Songklanakarin Journal of Science and Technology, 37(4), 409-415.

Hentschel, J., von Konrat, M., Söderström, L., Hagborg, A., Larraín, J.,Sukkharak, P., Uribe, J. & Zhang, L. (2015). Notes on early land plants today. 72. Infrageneric classification and new combinations, new names, new synonyms in Frullania (Marchantiophyta). Phytotaxa, 220(2), 127-142. DOI:

Sukkharak, P. (2015). A systematic monograph of the genus Thysananthus (Lejeuneaceae, Marchantiophyta). Phytotaxa, 193(1), 1-81. DOI:

Sukkharak, P., Pearaksa, P., Sarawan, A. & He, S. (2014). Bryophytes of Khao Kheow open zoo, Chonburi province. KKU Science Journal, 42(3), 571-578.

Sukkharak, P. & Khattiyawong, W. (2014). Bryophytes in Burapha University, Chonburi province. Thai Journal of Botany, 6(Special Issue), 1-4.

Sukkharak, P. & He, S. (2014).Bryophytes of beach forests in Chon Buri Province, Thailand. Songklanakarin Journal of Science and Technology, 36(5), 521-525.

Sukkharak, P., Kitlap, P., Likananonn, A. & He, S. (2014).A preliminary study ofbryophytes of the Khao Soi Dao wildlife sanctuary, Chanthaburi Province, Thailand. Songklanakarin Journal of Science and Technology, 36(5), 527-534.

Sukkharak, P. & Gradstein, S. R.(2014). A taxonomic revision of the genus Mastigolejeunea (Marchantiophyta: Lejeuneaceae). Nova Hedwigia, 99(3-4), 279-345. DOI:

Sukkharak, P. & Pikroapol, J. (2014). Bryophytes in Nong Tha Yu Arboretum, Chonburi province. Burapha Science Journal, (Special Volume), 115-122.

Sukkharak, P. & Gradstein, S. R. (2014). On the occurrence of asymmetrical underleaves associated with left-right symmetry in Mastigolejeunea, and the status of Mastigolejeunea undulata Gradst. & Grolle (Lejeuneaceae). Journal of Bryology, 36(2), 157-160. DOI:

Sukkharak, P. (2014). Studies on the genus Mastigolejeunea (Marchantiophyta: Lejeuneaceae): Mastigolejeunea gradsteinii Sukkharak sp. nov. Journal of Bryology, 36(1), 56-60. DOI:

Sukkharak, P. & Chantanaorrapint, S. (2014). Bryophyte studies in Thailand: past, present, and future. Cryptogamie, Bryologie, 35(1), 5-17. DOI:

Sukkharak, P. (2013). Jubula hutchinsiae subsp. javanica (Hepaticae: Jubulaceae): a genus and species new for Thailand. Polish Botanical Journal, 58(2), 585-587. DOI:

Ludwiczuk,A., Sukkharak, P., Gradstein, S. R., Asakawa, Y. & Glowniak, K. (2013). Chemical relationships between liverworts of the family Lejeuneaceae (Porellales, Jungermanniopsida). Natural Product Communications, 8(11), 1515-1518. DOI:

Sukkharak, P. & Asakawa, Y. (2013). Chemosystematics of the Thai liverwort Cheilolejeunea (Marchantiophyta, Lejeuneaceae). Natural Product Communications, 8(5), 565-566. DOI:

Sukkharak, P. & Gradstein, S. R. (2011). Response to “The New Guinean Thysananthus appendiculatus (Lejeuneaceae) discovered in the Western Ghats of India” by A. E. D. Daniels and R. D. A. Raja. Lindbergia, 34, 51-52.

Sukkharak, P., Gradstein, S. R. & Stech, M. (2011). Phylogeny, taxon circumscriptions and character evolution in the core Ptychanthoideae (Lejeuneaceae, Marchantiophyta). Taxon, 60(6), 1607-1622. DOI:

Sukkharak, P., Ludwiczuk, A., Asakawa, Y. & Gradstein, S. R. (2011). Studies on the genus Thysananthus (Marchantiophyta, Lejeuneaceae) 3. Terpenoid chemistry and chemotaxonomy of selected species of Thysananthus and Dendrolejeunea fruticosa. Cryptogamie, Bryologie, 32(3), 199-209. DOI:

Sukkharak, P. & Gradstein, S. R. (2010). On the occurrence of adnate underleaves and appendages in Thysananthus (Lejeuneaceae). Studies on the genus Thysananthus, 2. Journal of Bryology, 32(4), 308-310. DOI:

Sukkharak, P. & Gradstein, S. R. (2010). Studies on the genus Thysananthus (Marchantiophyta: Lejeuneaceae). 1. Thysananthus discretus Sukkharak et Gradst. sp. nov. Cryptogamie, Bryologie, 31(2), 113-119.

Sukkharak, P., Seelanan, T. & Lai, M. J. (2007). Liverwort diversity at the summit of Khoa Nan, Khao Nan National Park, Nakhon Si Thammarat province, Thailand. In H. Mohamed, B. B. Baki, A. Nasrulhaq-Boyce, P. K. Y. Lee (Eds), Bryology in the new millennium (pp. 49-55). Kuala Lumpur: University of Malaya.